diff --git a/model_signing/serialization/conftest.py b/model_signing/serialization/conftest.py
index addc9a6a..9dcf3af9 100644
--- a/model_signing/serialization/conftest.py
+++ b/model_signing/serialization/conftest.py
@@ -14,6 +14,8 @@
 
 """Test fixtures to share between tests. Not part of the public API."""
 
+import os
+import pathlib
 import pytest
 
 from model_signing.serialization import test_support
@@ -102,3 +104,12 @@ def deep_model_folder(tmp_path_factory):
         file.write_text(f"This is file f{i}.")
 
     return model_root
+
+@pytest.fixture
+def symlink_model_file(tmp_path_factory: pytest.TempPathFactory) -> pathlib.Path:
+    """A model with a symlink to a single file."""
+    source_file = tmp_path_factory.mktemp("source") / "file"
+    source_file.write_bytes(test_support.KNOWN_MODEL_TEXT)
+    symlink_file = tmp_path_factory.mktemp("symlink") / "file"
+    os.symlink(source_file.absolute(), symlink_file.absolute())
+    return symlink_file
diff --git a/model_signing/serialization/test_support.py b/model_signing/serialization/test_support.py
index 6d8aefaf..e3f45b04 100644
--- a/model_signing/serialization/test_support.py
+++ b/model_signing/serialization/test_support.py
@@ -39,6 +39,7 @@
     "empty_model_file",
     "empty_model_folder",
     "model_folder_with_empty_file",
+    "symlink_model_file",
 ]
 
 
diff --git a/model_signing/serialization/testdata/serialize_by_file/TestDigestSerializer/symlink_model_file b/model_signing/serialization/testdata/serialize_by_file/TestDigestSerializer/symlink_model_file
new file mode 100644
index 00000000..2ebc572a
--- /dev/null
+++ b/model_signing/serialization/testdata/serialize_by_file/TestDigestSerializer/symlink_model_file
@@ -0,0 +1 @@
+3aab065c7181a173b5dd9e9d32a9f79923440b413be1e1ffcdba26a7365f719b
diff --git a/model_signing/serialization/testdata/serialize_by_file/TestManifestSerializer/symlink_model_file b/model_signing/serialization/testdata/serialize_by_file/TestManifestSerializer/symlink_model_file
new file mode 100644
index 00000000..96d0bc82
--- /dev/null
+++ b/model_signing/serialization/testdata/serialize_by_file/TestManifestSerializer/symlink_model_file
@@ -0,0 +1 @@
+.:3aab065c7181a173b5dd9e9d32a9f79923440b413be1e1ffcdba26a7365f719b
diff --git a/model_signing/serialization/testdata/serialize_by_file_shard/TestDigestSerializer/symlink_model_file b/model_signing/serialization/testdata/serialize_by_file_shard/TestDigestSerializer/symlink_model_file
new file mode 100644
index 00000000..8ec1d11f
--- /dev/null
+++ b/model_signing/serialization/testdata/serialize_by_file_shard/TestDigestSerializer/symlink_model_file
@@ -0,0 +1 @@
+14aebf2e466ad30ef59ea6fce67de44dc133c673784bd543b45f75b8efc3d821
diff --git a/model_signing/serialization/testdata/serialize_by_file_shard/TestDigestSerializer/symlink_model_file_small_shards b/model_signing/serialization/testdata/serialize_by_file_shard/TestDigestSerializer/symlink_model_file_small_shards
new file mode 100644
index 00000000..7b4ad705
--- /dev/null
+++ b/model_signing/serialization/testdata/serialize_by_file_shard/TestDigestSerializer/symlink_model_file_small_shards
@@ -0,0 +1 @@
+beb3cbbd9d73133e85a102a3cbda2ef1dc2bc61e9323e32e576e4adb0571bf86
diff --git a/model_signing/serialization/testdata/serialize_by_file_shard/TestManifestSerializer/symlink_model_file b/model_signing/serialization/testdata/serialize_by_file_shard/TestManifestSerializer/symlink_model_file
new file mode 100644
index 00000000..6ef95249
--- /dev/null
+++ b/model_signing/serialization/testdata/serialize_by_file_shard/TestManifestSerializer/symlink_model_file
@@ -0,0 +1 @@
+.:0:22:3aab065c7181a173b5dd9e9d32a9f79923440b413be1e1ffcdba26a7365f719b
diff --git a/model_signing/serialization/testdata/serialize_by_file_shard/TestManifestSerializer/symlink_model_file_small_shards b/model_signing/serialization/testdata/serialize_by_file_shard/TestManifestSerializer/symlink_model_file_small_shards
new file mode 100644
index 00000000..3fdc9b2f
--- /dev/null
+++ b/model_signing/serialization/testdata/serialize_by_file_shard/TestManifestSerializer/symlink_model_file_small_shards
@@ -0,0 +1,3 @@
+.:0:8:a37010c994067764d86540bf479d93b4d0c3bb3955de7b61f951caf2fd0301b0
+.:8:16:bd762002a3528a27fb9a8822f822b949d3c9ab7e860af33039c9aa70ebbbe682
+.:16:22:a791e1e893ea4260c77475725101fb4cc6ad85f6340f21f10b239184e318cd21