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Code for analyzing RNA/protein compartmentalization, 3'UTR isoform expression, and fluorescence microscopy data in healthy and ALS patient-derived cell lines.

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ALS Compartmentalization

Code for manuscript: [Link TBD](Link TBD)

Dependencies

R (Version 4.3.1)

To install required libraries, the load_libraries.r script can be invoked as follows:

RScript load_libraries.r

Code Organization

Code for 3'UTR annotations and quantifications can be found at: 3utr_annot and 3utr_quant

Code for fluorescence microscopy processing can be found at: ALS_MN_image_analysis

The codebase is organized according to source dataset:

  1. MN_Diff (motor neuron differentiation time course)
  2. Astrocytes
  3. siRNA knockdown: targeting SFPQ, FUS, TDP-43
  4. VCP knock-in model with ML240 treatment
  5. Microscopy images of motor neurons (Stressed_MN)

and perform the following analysis:

Analysis Script Description Figure(s)
RNA Compartmentalization 1_GetRNAMass.r Analyzes cytoplasmic and nuclear RNA mass at each stage of MN differentiation. Fig 2A
2_Plot_PUD_GE_Nuc_Cyto_Cor.r Analyzes correlation between nuclear and cytoplasmic gene expression or 3'UTR usage profiles. Fig 2C
3_GetGenomicFeaturesDistribution.r Analyzes fraction of mapped reads to genomic features (Introns, Exons, 3'UTRs, 5'UTRs). Fig 2D
Protein Compartmentalization 1_PlotProtExp.r Plots protein relative abundance (fold-change compared to iPSC stage) for NEFL. Fig 2B
2_Plot_Prot_Nuc_Cyto_Cor.r Analyzes correlation between nuclear and cytoplasmic protein abundance profiles. Fig 2C
3'UTR Analysis 1_Identify3UTRShifts.r Identifies significant shifts in relative expression of 3'UTR isoforms across conditions using Fisher's test. -
2_Process3UTRShifts.r Processes results from the Fisher's test. -
3_Plot3UTRShifts.r Generates volcano and barplots of 3'UTR isoform shifts; performs GO enrichment and pathway comparison. Fig 2E, 2F, S2–S5
4_PlotPUDBarplots.r Bar plots for specific genes illustrating 3'UTR isoform expression across conditions. Fig 2G, 2H, S1B
5_PlotUTRLength.r Generates barplots for overall 3'UTR lengths across conditions. Fig S1A
6_qPCR.r Analyzes qPCR results for ENAH 3'UTR isoforms. Fig 2I
RBP (CLIP-Seq) and m6A Analysis 1_CLIPGlobalRegWelchTtest.r Identifies global RBP regulators of 3'UTR usage using Welch’s t-test; generates heatmaps and lineplots. Fig 4A, 4B
2_CLIPEnrichmentFisher.r Identifies enrichment of RBP binding comparing shifted vs non-shifted 3'UTRs; heatmaps of enrichment/depletion. Fig S7
3_FindRBPBindingSites.r Plots density of RBP binding in specified 3'UTRs. Fig 4D
4_FindRNAModSites.r Plots density of m6A modification in specified 3'UTRs. Fig 4E
5_Compare3UTR.r Compares RNA modification in shifting vs stable 3'UTRs. Fig S7C
6_FindTDP43-m6aOverlap.r Finds overlap between m6A and TDP-43 binding by mutual information calculation. Fig 4F
7_PlotM6aActivity.r Plots m6A activity in iPSC and MN cell lines. Fig 4G
Microscopy Analysis 1_Radius.r Generates plot for radius of proximal or distal neurites in VCP and WT samples. Fig 3D
2_RBPLoc.r Generates plot for relative intensity of TDP-43 in cytoplasm, proximal, or distal neurites in VCP and WT samples. Fig 4H

Usage

The config file config.yml is used to specify location of input data and input/output directories.

Each script can then be invoked, for example, as follows:

RScript ./scripts/MN_Diff/3UTR_Analysis/1_Identify3UTRShifts.r config.yml

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Code for analyzing RNA/protein compartmentalization, 3'UTR isoform expression, and fluorescence microscopy data in healthy and ALS patient-derived cell lines.

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